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ACM
Tropical Field Research in Costa Rica
A
listing of topics and abstracts from Tropical Field Research program
participants, arranged by subject area. Copies of these papers and
projects are kept on file at the ACM office in Costa Rica for
use by ACM program participants only.
Each
entry includes: student's name, topic, advisor's name, student's
college, year of participation, length of paper, abstract.
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Botany
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Barber-Gormley,
Peregrine, “Ethnobotany and the study of optimal natural ecological
conditions of an endangered cycad, Zamia skinneri, for the
enhancement of future cultivation techniques.” Adv. Rafael Ocampos.
Colorado. 1999. 77 pp.
The
environmental conditions of two wild populations of the Costa Rican
cycad, Zamia skinneri, were studied in Baltimore, Matina
county, Limón province, Costa Rica. Slope and soil conditions were
analyzed to determine the effects on population abundance. The rise
and the run of slopes was measured in a total of 62 plots containing
the two populations. Higher mean population densities were found
on moderate to steep slopes, than on flatter slopes. Both physical
and chemical soil conditions were analyzed from two composite soil
samples taken separately from the two population sites. Soil conditions
showed considerable differences between the two populations. The
information found in this study, in conjunction with previous studies,
seems to show that Z. skinneri prefers sands and sandy loams
over soils with high clay contents. However, no definitive statements
about the influence of chemical composition correlating with changes
in population abundance, can be made. The information collected
in this study, will aid in the artificial cultivation of this endangered
ornamental plant. An ethnobotanical study was conducted in the Cabecar
community of the Chirripo indigenous reserve in order to assess
current knowledge of Zamia skinneri, and increase local awareness.
The study determined that the majority of the population can identify
the plant, but lack knowledge of its medicinal use. Out of four
families interviewed, one used the sap of this plant to speed the
healing of wounds. Also, out of 21 elementary school students, three
stated that their families used the plant for wound healing, and
one of those stated that it could also be used to eliminate the
parasitic skin larva, Papalomoyo (Leish maniases).
Bay, Robin, “Comparison of plant species in three differing
sucessional communities of the tropical dry forest in Palo Verde
National Park, Guanacaste.” Adv. Michael McCoy. Colorado College.
1999. 15pp.
This study looked at the secessional process of the tropical dry
forest in Palo Verde National Park, Guanacaste during March and
April. Differences in tree species composition, frequency, diversity
and percent cover of the understory were compared between three
communities of tropical dry forest that exhibited varying degrees
of disturbance by cattle grazing. The first (P1) was not grazed
between 1980 - 86, but now is grazed intermittently. The other two
(P2 and P3) were grazed in the past but have not been for the last
twenty years, although P3 burned in 1989. All three areas are bordered
by mature relatively undisturbed forest.The plant species diversity
was found to be significantly greater in the forests than the pastures
and also greater at site 2 than either of the others. The percent
cover of woody species in the understory exhibited the same trends
as well, although in Site 2, percent cover was the same in the forest
and pasture suggesting greater recovery at this site. Supporting
this, the species composition of the two habitats at Site 2 showed
greater similarity than at sites 1 and 3. The frequencies of the
most common species in each site were compared between sites and
habitats. The frequencies of those species most commonly foraged
by cattle were also compared.
Blank, Vicki L., “A Comparison of Forest Structure and Tree
Species Composition in Three Successional Sites of Cloud Forest
in Monteverde, Costa Rica.” Adv. M. Sc. Dora Ingrid Rivera. Grinnell.
1999. 17pp.
This study analyzes the changes in tree species composition of Monteverde?s
neotropical cloud forest as areas of forest recover from human disturbance.
Data on tree species diversity and abundance were collected in a
primary forest and secondary forests of approximately eight and
forty years. With respect to DBH, height, and density of individuals,
the forest structure of the 8-year forest was simple in comparison
to those of the 40-year and primary forests, while the 40-year and
primary forests were similar in structure. However, with respect
to species richness, diversity indices, and levels of oligarchy,
the species composition of the primary forest was more complex than
those of both the secondary forests. Indices of similarity between
forest types also support the conclusion that forty years is not
sufficient for a secondary forest to regain the biodiversity present
in a primary cloud forest.
Brenner, Jacob, “Branch attachment and survival of fallen
epiphytic bromeliads (Bromeliaceae), Cordillera de Talamanca, Costa
Rica.” Adv. Christopher Vaughan. Lawrence University. 2000.
I studied survival of fallen epiphytic bromeliads in a neotropical
montane rain forest on the Dúrika Biological Reserve, Puntarenas
Province, Costa Rica during March and April, 2000. The objective
of this study was to determine the influence of host branch attachment
on survival of these bromeliads on the forest floor. I sampled,
identified and characterized 1,496 plants in ten species. Results
showed significantly different survival percentages across five
attachment classes: large branch, small branch, twig or bark, detached
with roots and detached without roots. Survival depended on attachment,
decreasing with branch size. I discuss the principal factor as branch
substrate security.
Candelaria, Karin, "The Effects of Habitat on Plant Species
Composition in Páramo Las Vueltas, Cordillera de Talamanca, Costa
Rica." Adv. Carlos Valerio. Knox. 1991. 28pp.
The effects of habitat on the composition of plant species were
analyzed within various plant communities of Páramo Las Vueltas,
in the Cordillera de Talamanca, Costa Rica during March and April,
1991. Special consideration was given to soil composition and climatic
factors within the area. The páramo is loosely defined as the type
of vegetation which occurs between the timberline and the snow line
in the tropic belt. Five sites were chosen on the páramo in order
to sample plant community types. Within each site, five quadrants
were surveyed, each of twenty-five square meters. Frequency, density,
and cover were measured for each plant species. An importance value
was assigned to each species based on the sum of the relative values
of these parameters in order to have an idea of the dominance of
the species within the area. The depth of the top layer of soil
was also measured for each site. Soil samples were taken and analyzed
for the content of organic and inorganic material. From these tests,
it was found that the species composition of plant communities is
affected less by variations of edaphic conditions and more by local
differences in climate, topography, wind, sunlight, and inclination
of slope.
Crandall,
Suzannah, "Fungal Infestation and pest Damage on Theobroma
cacao Pods in a Premontane Wet Forest zone of Costa Rica."
Adv. Christopher Vaughan. Beloit. 2003.
This
study will document the occurrence of fungal infestation and pest
damage to Theobroma cacao pods over a period of approximately five
months as the pods mature. Approximately 1000 pods will be sampled
from different heights and tree clones to provide a detailed record
of specific causes of damage and to quantify pod mortality. This
information will be used to determine options for controling damage
in the future.
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Davidson,
Anna, "Habitat Preferences and Comparative Sugar Analysis in
Leaves of Selected Symplastic Phloem Loading Species of Bignoniaceae
and Acanthaceae in the Humid Forest of Sarapiquí and the
Dry Forest of Santa Rosa, Costa Rica." Advisor: Walter Marín. Ripon
College. 2005. 21pp.
Phloem loading is the starting point of long distance nutrient transport.
There are two types of phloem loading. Symplastic loading occurs
through the plasmodesmata-connected cytosol of cells. Apoplastic
loading requires at least one extracellular step through cell walls.
Symplastic phloem loaders have unique characteristics that distinguish
them from apoplastic phloem loaders such as intermediary cells (a
specialized companion cell), highly branched plasmodesmata more
so on the bundle sheath cell, and they translocate raffinose and
stachose unlike apoplastic loaders that transport only sucrose.
It has been observed that intermediary cells have evolved at least
five different times. What are the adaptive advantages of symplastic
phloem loading? Why are there two mechanisms of phloem loading?
To help answer this question, a floristic analysis was done and
illustrated that symplastic phloem loaders are most prevalent in
tropical environments. What are the characteristics of a tropical
environment that might be involved in adapting to loading symplastically?
The objective of this research is to compare the habitat preferences
of two symplastically phloem loading families, Bignoniaceae and
Acanthaceae in two extremely different environments (dry tropical
forest and wet tropical forest) within Costa Rica, a country that
significantly represents a tropical environment. This research may
help us to narrow down characteristics of tropical environments
that might have aided in the evolution of symplastic phloem loading.
It was found that there is almost equal diversity of Bignoniaceae
species in both the dry forest and the wet forest. However, there
is a higher density of Bignoniaceae in the dry forest. Acanthaceae
has a higher number of species in La Selva versus the dry forest
of Santa Rosa. Families as a whole do not seem to prefer one environment
to another, their preference is at the species level. Leaf material
of each species was taken and ground with ethanol. The sugars of
these samples will be later analyzed at Cornell University using
HPLC to identify the sugars, and the amount of sugar between species
of the same family that occur in these two different environments.
Driessen,
Brad, "Savanna Succession in the Premontane Moist Forest of
Turrúcares, Alajuela Province of Costa Rica." Adv. Alvaro Castaing.
Lawrence. 1992. 58pp.
An
ecological study of relic forest sites along the Tizate River located
near Turrúcares, Alajuela undergoing a plant succession to Savanna
forest. Frequency, density, and dominance values of trees were found
to determine the importance of species within the sites. As a result
of the annual burning of pastures, substitution of larger, perhaps
native tree species by smaller fire resistant species was found
to be occurring. This is leading to the degradation of soils and
conditions of the water shed. The researcher offers some appropriate
techniques to do Social Forestry to increase the productivity of
the land to reverse the process of savanna formation.
Gerstner,
Brett Justin, "Distribution and Diversity of Epiphytic Ferns
on Tree Bases in Costa Rican Montane Cloud Forests." Advisor: Maarten
Kappelle. Lawrence University. 2005. 24pp.
This study assessed the diversity and distribution of epiphytic
ferns on tree bases in the upper montane oak dominated cloud forests
near San Gerardo de Dota, in the Talamanca mountains of Costa Rica.
Epiphytic ferns dwelling on the bases of 150 oak and non-oak tree
stems were sampled in three ecologically different forest stands:
one in a mature, old-growth forest on the drier Pacific slope, and
two in Successional forests inhabiting both the drier Pacific slope
and wetter Atlantic slope. In total, 206 fern individuals were recorded,
distributed over 16 species in seven genera. Using different diversity
indices, the wetter Atlantic stand turned out to be most species
rich, being 1.6 to 1.7 times richer than the drier Pacific stands.
Atmospheric humidity seems to be the main factor determining species
diversity. Two distinct species groups were found in two forest
stands. Increased stem diameter seemed to only slightly enhance
the diversity of epiphytic ferns on stem bases. Four epiphytic fern
species seem to have a preference for specific tree stem diameter
classes, though further study is needed to analyze this pattern.
It is concluded that humidity seems to be much more important than
successional age as a key factor in determining fern species diversity.
Thus, it is postulated that on the long term climate change (in
this case, changing precipitation patterns and cloud cover) may
locally have a greater impact on epiphytic fern species diversity
than forest clearing and subsequent natural recovery.
Gordon,
Sarah, "20 Medicinal Plants Used by the Bribri Indians in Coroma,
Talamanca." Adv. Juan Diego Alfaro. Colorado. 1992. 48pp.
This is an ethnobotanical study of the medicinal plants used by
the Bribri Indians in Coroma, Talamanca. The study site is the Bosque
Comunal, a 3,000 ha. portion of virgin tropical premontane wet forest
used by the Bribri to practice and preserve their traditional ways.
I identified 20 medicinal plants used by the Bribri that are found
in the Bosque Comunal and in the Talamanca Valley at the base of
the area. Information on how the Bribris use these medicinal plants,
and how they are tied into Bribri mythology is discussed. In addition,
the significance of the Bosque Comunal in light of the constant
pressure of deforestation in the Talamanca Valley is included. Finally,
suggestions for future projects are discussed.
Galey,
Kelly, "Physical and Phytosociological Observations on the Natural
Regeneration of Cordia alliodora, Upala, Northern Costa Rica."
Adv. Alvaro Castaing. Macalester. 1998. 24pp.
Over the months of March and April of 1998, a study of the natural
regeneration of Cordia alliodora, commonly known as laurel,
was conducted in Pueblo Nuevo de Bijagua, near Upala in northern
Costa Rica. The main objective was to compare the growth of laurel
in different stages of progression. Biometrical data was collected
on seedlings and on mature trees. The distance between both seedlings
and mature trees to the nearest mature tree was measured. Health
was measured in seedlings as a ratio of diseased leaves to total
leaves. The presence or absence of nodules in seedlings was recorded.
The only significant difference found between the two populations
was the density which was higher in primarily seedling populations.
A more diversified plant population was found in the primarily seedling
population. Growth of seedlings, in terms of height and diameter,
were significantly related to their distance from the nearest laurel
tree by regression analysis. The presence of nodules was significantly
related to the diameter and height of the seedling by regression
analysis, but had no significant relation to the health of the seedling.
There was also a significant correlation between the circumference
at breast height of mature trees and the proximity of the nearest
mature tree. While the growth of seedlings found growing within
the mature population was not significantly different from those
in seedling populations, seedling density was affected. Additionally,
both seedlings and mature trees growth decreased as the distance
to the nearest neighboring tree decreased.
Groome, Sarah, "Arrabidaea chica (Bignoniaceae): an
ethnobotanical study of its biology, domestication potential and
uses by two indigenous groups, the Bribri and Cabécar of Costa Rica."
Adv. Rafael Ocampo. Colorado. 1998. 29pp.
The first formal study of Arrabidaea chica (Bignoniaceae)
in Costa Rica was conducted during March and April, 1998, in the
Cantón of Matina, Limón, Costa Rica. The first half of the study
focused on learning about the biology and ecology of the vine, by
establishing parcels in different forest environments located in
the farm Bougainvilla S.A. in La Esperanza de Matina, Limón. Cuttings
of the plant were taken and grown in artificial conditions to obtain
an idea of its domestication ability by asexual reproduction and
to observe its potential as a non-timber forest product. The second
half of the study focused on investigating the knowledge and uses
of this vine by two indigenous groups, the Bribri and Cabécar speaking
peoples of Costa Rica, by the use of informal interviews. The most
common use of Arrabidaea chica for the Bribri is as a red dye used
to color fibers to make baskets and other forms of craftsmanship;
uses by the Cabécar remain ambiguous. This research provides preliminary
data that the plant is found abundantly in forest areas with more
light, in various soil types, has important usefulness for the Bribri
of Costa Rica and has economic potential as a non-timber forest
product, medicinal plant and natural colorant.
Hart,
Georgia M., "Entomophilous pollination of coffea arabica
(Rubiaceae) and its pollination interations with neighboring wild
confamilials." Adv. Luis Diego Gómez. Grinnell. 2003.
This
project will investigate the reproductive consequences of insect
pollination for Coffea arabica and its interactions with
other Rubiaceae in forest surrounding coffee plantations near San
Vito in the southern Talamanca range. Insect pollinators will be
identified and the percentage of pollination of C. arabica
through insect vectors and other means such as wind pollination
and autogamy will be calculated. The effects of coffee pollen and
pollinators will be tested on neighboring confamilials.
Hayford,
Kyla, “A comparison of understory plants in three successional
cloud forests in Monteverde, Costa Rica.” Adv. Dora Ingrid Rivera.
Macalester. 1999. 20pp.
Succession is the natural change in species composition after a
disturbance. Forests can usually recover from natural disturbances
yet human-derived disturbances, such as deforestation, may be too
overwhelming to recover from. Generally, a mature forest will have
greater diversity. I studied the effects of deforestation by performing
a quantitative inventory of the species in the understory at the
Sky Trek reserve in the Monteverde Cloud Forest. 8 year, 40 year
and primary forests were compared and analyzed according to the
Shannon-Weiner Diversity Index, overall density, family richness,
and percent similarity. The preliminary study supported the Intermediate
Disturbance Theory in which the family richness and diversity were
highest in the 40 year forest and then decreased in the climax primary
forest. According to the Shannon-Weiner Diversity Indices, the 40
year forest has recovered to the level of a primary forest. However,
the percent similarity, family richness, and overall density suggest
that the secondary forests have not fully recovered. The successional
pathways are characterized by five major families. Thelypteridaceae
and Urticaceae are colonizing families where as Dryopteridaceae
is a climax family. Both Rubiaceae and Piperaceae are abundant in
all stages of succession.
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Johnson,
Kaia, “The phenotypical characterization and harvesting of Smilax
(Cuculmeca) in the tropical forest of Limón, Costa Rica.” St. Olaf.
2000. 30pp.
A study of Smilax spp. (Cuculmeca) was conducted during March
and April, 2000, in the county of Matina, Limón, Costa Rica. Research
took place in secondary tropical forest in the Agroecological Garden
Bougainvillea S.A. Investigation involved the phenotypical characteristics,
domestication and conservation of Cuculmeca. Thirty-nine adult plants
were found, measured and observed. Samples of flowers and fruits
were taken; and seeds were counted and planted. Pieces of rhizome
containing shoots were planted in parcels in the tropical forest.
Ethnobotany determined the method of harvest and indicated that
three traditional classes of Cuculmeca exist: blanca, roja and morada.
Information from this investigation will provide preliminary data
which may be used in future studies on Cuculmeca.
Kashi,
Natalie, "Cyclanthaceae and Weevils." Adv. Luis Diego
Gómez. Colorado. 2003.
This
study will research the pollination syndrome of one or two species
of the Carludovicoideae subfamily of Cyclanthaceae. Evidence of
the pollination pattern of Cyclanthaceae will provide more information
on the phylogeny of Cyclanthaceae and the relationship between ancient
insects and plants.
Kroese, Jesse, "Gall richness in the Paramo and high altitude
oak forest of Cerro de la Muerte with an emphasis on Melastomataceae."
Adv. Paul Hanson. Macalester. 1998. 12pp.
Variations in galling species richness and galling patterns in tropical
plant families are two aspects of gall ecology that have not been
extensively studied. This investigation addressed these questions
through two sub-studies. Study A: Two hypotheses which attempt to
explain the variation in species richness of gall-formers in different
life zones were tested in three Costa Rican life zones: the montane
rain forest (high altitude oak forest), the Paramo and the wet tropical
forest. The number of galling species found over time was recorded
for transects 2 x 50 meters in dimension. Galling richness in oak
forest and wet tropical forest was not significantly different (
= 0.738, df = 1, p = 0.384), while there were highly significant
differences in the Paramo-Oak forest comparison ( = 26.31, df =
1, p < 0.0001 ) and Paramo-Cańo Palma comparison ( = 12.14, df =
1, p < 0.000493). These results did not conclusively support or
refute the two hypotheses in question. Study B: Seven species of
Melastomataceae were surveyed for galling patterns in the Paramo
and high altitude oak forest. Six of the species contained galls;
nine types of galls were present, with an average of 0.806 galls
per plant. Galls per plant frequencies did not closely follow the
normal (Gaussian) distribution.
Kurth,
Valerie, "A Comparison of the Natural Regeneration of Carapa
guianensis in a Cleared Pasture and in a Primary Forest in the
Northern Zone of Costa Rica (Upala)." Adv. Alvaro Castaing. Macalester.
1998. 14pp.
The
natural regeneration capabilities of the tree species caobilla (Carapa
guianensis) were studied on a cattle farm in Pueblo Nuevo de
Bijagua (canton: Upala; province: Alajuela, Costa Rica) during the
months of March and April, 1998. The purpose of the study was to
compare the regeneration of caobilla in a cleared pasture and in
a primary forest. The diameter and height of seedlings growing beneath
mature trees was measured and their herbivory status was observed.
The distance between mature trees and seedlings was measured. Seedling
diameters and heights were greater in the forest than the pasture
(F = 5.86; df = 1, 962; p = 0.0157; F = 123.63; df = 2, 745; p <
0.00009, respectively). Percent herbivory was greater in the pasture
than the forest (F = 5.86; df = 1, 20; p = 0.0251) and the mean
distance between mature trees and seedlings was greater in the forest
(F = 13.84; df = 2, 396; p < 0.00009). These results suggest vertebrate
predators are responsible for seed dispersal in the forest while
pasture seedlings tend to develop where they land beneath the mature
tree. Forest seedlings also develop faster and suffer less from
herbivory than pasture seedlings.
Kusmin, Victor N., "Dracontium sp. (Araceae): Un estudio
etnobotánico con los indígenas cabécar de Costa Rica." Adv.
Rafael Ocampo. 1997.
An unidentified species of the little studied genera Dracontium
(Araceae) was studied formally for the first time in the
northern part of the Talamanca region, Limon. The first half of
the study focused on a population growing within the agroecological
system, of cacao (Theobroma cacao), timber, native and other
domesticated species, present at the farm Bougainvilla S.A. in La
Esperańza de Matina, Limon. An attempt was made to identify the
species, and gain a knowledge of the biology and ecology of the
plant in respect to its potential as a non timber forest product.
To this end, collections were made, and various measurments taken
from a range of plant sizes and compared. The second half of the
study focused on investigating what remains of the knowledge of
the traditional and historical uses of hombrón by the indigenous
Cabécar speaking peoples. Various formal and informal interviews
were conducted, in Spanish, and recorded. The most commonly known
use of this genus is its function as a remedy for the venemos terciopelo
(Bothrops atrox). While very little remains of the Cabécar
knowledge of the traditional uses of this, one of the largest, species
of Dracontium, it does posses economic potential as a non
timber forest product, in respect to a tropical agroecological system.
Lila, Rasa, "Dracontium gigas (Araceae): Ethnobotonical
Study and Enrichment of the Tropical Forests with a Medicinal Plant
Used by the Bribri and Cabécar Indigenous People of Costa Rica."
Adv. Rafael Ocampo. Colorado. 1998. 53pp.
In the Spring of 1997, a formal study on an herbaceous plant Dracontium
gigas (Araceae) was conducted for the first time in the northern
part of the Talamanca region of Limón, Costa Rica. This plant is
used medicinally by the Bribri and Cabécar Indigenous people of
Costa Rica to treat venomous snake bites. The first study focused
on the investigation of the biological aspects of Dracontium. There
was a need for further research on its ecological limits, taxonomy,
life cycles, history and use by the Indigenous peoples. The present
study focused on these aspects as well as the planting of Dracontium
in the forest to enrich it, and preparing for future investigations.
If Dracontium is shown to be an important forest plant resource,
it could contribute to a group of plants called nontimber forest
products. These plants offer an economic alternative to cutting
down the Tropical Forests. The preliminary results found the studied
Dracontium plants to be the species D. gigas. The findings
supported that Dracontium thrives in disturbed environments and
agroecological systems more than primary forest. It was also found
to be an extremely important forest plant resource for the Indigenous
Bribris and Cabécar to cure venomous snake bites.
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Lobato,
Tonya, "A Field Guide: An Introduction to the Vegetation of
the Volcán Cacao Biological Station" Adv. Eric Olson. Lawrence University.
1997. 27pp.
The plant and tree species listed in this guide were collected from
the 800 meter trail through the pre-montane tropical moist forest
accessing the Volcán Cacao Biological Station. They represent some
of the most common or conspicuous species growing along the trail,
and provide only a small sample of the astounding plant diversity
found there. This guide was written in order to provide visitors
and students of Volcán Cacao a means of familiarizing themselves
with the vegetation whether it be for academic purposes or simply
individual interest. As much information as possible was recorded
about each species. All listed species include a basic pictoral
representation along with a description pointing out distinguishing
characteristics of the plant or tree. The map included in the beginning
of the guide points out specific trees along the trail with numbers
corresponding to the numbered listings in the book. For the species
listed on the map, depending on the abilities of the navigator,
identification can be quite easy; the others, however, may require
a bit more effort. Careful observation, along with the information
provided in the book, should enable most individuals to correctly
match the remaining species with their live counterparts. Each species
is listed under its appropriate family; the families are arranged
in alphebetical order.
Logback,
Judy, "A Study of Successional Process in 'Openings' in the
Premontane Tropical Rain Forest of San Bosco." Adv. Alvaro Castaing.
Beloit. 1993. 20pp.
The ecological process of tropical rain forest succession in forest
'openings' was studied by comparing characteristics in the stages
of pioneer, early secondary, late secondary and virgin forest located
in the Cordilleras of Costa Rica. The microclimates and biodiversity,
frequency, and density of scientific families were investigated,
in premontane forest successional sites of the San Bosco Region.
Lucas,
Rachel, "An Evaluation of Nectar Production and Quality of Heliconia
tortuosa Griggs (Heliconiaceae) with Ambient Temperature." Advisor:
Luis Diego Gómez. Lawrence University. 2005. 50pp.
This research evaluates the interaction between environmental factors
that may cause variations in nectar production and quality. Understanding
the causes of variation in nectar production may help with further
insight into co-specfic relationships between hummingbirds and flowers.
Las Cruces Biological Station in San Vito, Coto Brus, Costa Rica
provided an abundance of heliconia plants to study during March
and April of 2005. Data collection included temperature, nectar
volume, nectar concentration and pH every two and a half hours from
0630 hr to 1600 hr. In addition, weather data was collected from
a weather station at Las Cruces; these environmental factors were
used to analyze ambient factors in relation to nectar production
and quality. Nectar production and total sugars concentration has
a significant relation with time. Temperature and other ambient
factors were found to have very little effect on nectar production
and concentration.
Meisner-Bogdahn, Hanna, "Effects of Climate Change in Monteverde
Cloud Forest on Survival, Growth, Reproductive Success of Platystele
microtatantha: A Proposal." Adv. Karen Masters. Colorado.
2003.
This
study will address how climate change during the dry season in Monteverde
and more specifically, the decrease in dry season mist frecuency,
affects survival, growth and reproductive success of the Platystele
microtatantha orchid species. It will also investigate the water
conservation strategies used by P. microtatantha under conditions
of decreased mist frequency compared with conditions of more consistent
mist.
Okrent,
Rachel, “Distribution and abundance of Smilax spp. (Cuculmeca
and Zarzaparrilla) in three permanent plots in secondary growth
tropical humid forest on the Caribbean coast of Costa Rica..” Adv.
Rafael Ocampo. Grinnell. 2000. 20pp.
The distribution and abundance of Smilax spp. (Cuculmeca
and Zarzaparrilla) was investigated in three permanent plots in
secondary growth tropical humid forest in Limón, Costa Rica. These
medicinal plants, thought to be in danger of extinction, demonstrated
a stable population structure at the study site, Garden Bougainvillea
S.A. Plants were categorized into the following three development
states: regeneration, juvenile, and adult. The census revealed 600
plants of the common name Cuculmeca and 94 of Zarzaparrilla. There
exist 565 regenerates, 70 juveniles and 56 adults. Ethnobotanical
investigation, performed in conjunction with the ecological study,
revealed differences in the knowledge regarding Cuculmeca between
indigenous and non-indigenous people of the area.
Oyama,
Ryan, "Characterization of a Premontane Wet Forest Dominated
by quercus seemanii in Costa Rica." Adv. Dr. Alvaro Castaing.
Lawrence. 1995. 16pp. This study attempts to characterize a Tropical
Premontane Wet Forest dominated by Oak (Quercus seemanii).
This forest was located at a relatively low altitude on a ridge
near El Carmen de Parrita, Costa Rica. Plant samples were collected
along a one kilometer transect and identified to at least the family
level. Profile measurements of the trees along the transect were
also made. From this data, a picture of the community composition
was obtained. The forest was found to be highly diverse and transitional
to a Lower Montane Wet Forest due to a steep climatic gradient that
exists along the ridge. The regeneration of Oak seedlings was also
studied and no significant correlation to light was found.
Pearson, Duane, "Population Dynamics of the Cedro Macho Tree,
Carapa guianensis in the Tiskita Biological Reserve." Adv.
Alvaro Castaing. Lake Forest. 1992. 12pp.
Population dynamics of a ridge top population of Carapa guianensis
was studied in the Tiskita biological reserve. Basal area, seed
size and dispersal patterns, biomass production, germination success
rates, and seedling dispersal were all studied. Basal area for this
ridge top community was 7.66 m2/ha. Average seed weight was 74.2g.
Leaf Litter production for the forest as a whole was 22.6g. dry
weight per m2 per week. 21% (4.75g.) of this biomass weight was
contributed by C. guianensis. Dispersal patterns fell into
the preestablished norms for this species: the number of seeds present
on the ground increases as distance from the trunk of the parent
tree increases. 75.5% of these seeds germinate. The highest concentration
of yearling seedlings (smaller than 1.5 meters altitude) was found
within one meter of the parent tree. The hypothesized reason for
this pattern of seedling dispersal is an increase in soil humidity
and organic material next to the trunk. This pattern of seedling
dispersal had no apparent effect on the dispersal patterns of the
adult and juvenile members of this species. The results of this
study were compared to the results of two studies that were done
with C. guianensis in its climax habitat: moist, poorly drained
soils. Population density was much lower on the well drained slopes
of the Tiskita reserve. Seed size was significantly increased, double
what had previously been reported. Seed dispersal patterns were
not changed by differences in terrain.
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Peterson,
Snow Brook, “Differences in flower characteristics and insect
visitor composition for two species of Ageratina (Asteraceae)
on Cerro de la Muerte, Costa Rica.” Adv. Paul Hanson. Grinnell.
2000. 18 pp.
This study investigated differences in insect visitor rates and
flower characteristics between two superficially similar species
of the genus Ageratina (Asteraceae) -- A. ixiocladon
and A. reticulifera. The study was conducted during March
and April of 2000 between the towns of Jaular and Villa Mills, Costa
Rica, in a tropical montane rainforest life-zone. I made a series
of observations of different individuals of each plant species and
found that A. ixiocladon was more frequently visited by bumblebees
than A. reticulifera, while A. reticulifera was more
frequently visited by tachinid flies. Both types were visited by
equal numbers of total visitors and syrphid flies. The corolla tubes
of A. ixiocladon are significantly longer than those of A.
reticulifera, and their flower heads also average a higher number
of florets. The differences in visitor composition, which may be
correlated to the differences in flower characteristics, contradict
the theory of similar looking Asteraceae acting as generalists in
the insects they attract (i.e. the insects do not distinguish between
species).
Reed,
John, "The Pollination Biology of Conostegia xalapensis
(Melastomataceae)." Adv. Sally Bianco. Grinnell. 1990. 17pp.
During the months of March and April 1990 the pollinated biology
of Conostegia xalapensis (Melastomaceae) was studied
in the area of Santa Elena, Costa Rica. The hermaphroditic flowers
are physically open to pollination for one day (0600-2100) although
the stigmata remain receptive to pollen for another 24 hrs. C. xalapensis
was found to be an outcrosser with a 99.6% fruit set from manually
cross-pollinated flowers vs. a 4.5% fruit set from manually self-pollinated
flowers. The principal pollinators were found to be Melipona
beechei and M. fasciata (diurnally active stingless bees
of the family Apidae).
Rickards,
Anna, "Abundance, Diversity, and Distribution of Epiphytic Plants
on Fallen Trees in a Tropical Premontane Rainforest." Adv. Dr. Walter
Marín, UCR. Oberlin College. 2006.
This study assesses the diversity, distribution, and abundance of
all the epiphytic plants found on three different fallen trees in
the tropical and premontane rain forest (wet forest) of the Organization
of Tropical Studies' (OTS) La Selva Biological Field Station and
Reserve. All the epiphytes found on the tree were recorded along
with their height and position. Representative species were collected
for the purposes of identification and collecting height and weight
measurements. Data has been organized in various ways to obtain
a full picture of the epiphytic cover of each tree, including both
Plant Height and Plant Weight vs. Circumference of Phorophyte, Distribution
of Common Families among Branch Size Categories, and a map of all
the epiphytes found on each tree indicated by family.
Ridley,
Caroline, “Cecidomyiid galls on Peperomia species in
a tropical montane cloud forest on Cerro de la Muerte, Costa Rica.”
Adv. Paul Hanson. Grinnell. 2000. 16pp.
During the months of March and April 2000, I investigated a little
known gall type induced by members of the family Cecidomyiidae,
also known as the gall midges, on plants of the genus Peperomia
on Cerro de la Muerte, Costa Rica. This study was a preliminary
examination of the morphology of these galls, as well as a general
characterization of the Peperomia plants which are used as
hosts. Results indicate that one species of cecidomyiid is responsible
for inducing morphologically similar galls on three of eleven species
of Peperomia found within the study site. Variation in gall
color seems to be due to differences in gall age while variation
in gall size appears to be attributable to multiple factors including
host species, age, and the number of larvae within the gall.
Schoeb,
Brent, "An Analysis of Primary and Second Growth Montane Rain
Forest Near La Trinidad de Dota, Costa Rica." Adv. Carlos Valerio.
Lawrence. 1991. 19pp.
In
this study a series of plots within 40 ha. of primary montane forest
and 30 ha. of second growth forest were analyzed. The secondary
forest is the result of five years growth after this section of
land was clearcut, with the exception of several large trees which
remain today. The area for this study is located in the Cordillera
de Talamanca, Costa Rica, at an elevation of ca. 2700 m. 25 m2 plots
were used within both forest types to identify the woody species
present (individuals d.b.h. > 20 mm.) and to map these individuals.
A direct replacement of the dominant species in the primary forest
was observed in the secondary forest, presumably due to remnant
seed trees present in the latter. The notable exception to this
was the lack of Quercus spp. in the secondary forest, possibly due
to predation on acorns of these species. In addition, data regarding
the basal area, height, and diameter at breast height for each forest
type are presented here.
Stavina,
Nina, “Ethnofarmacology and Methods of Reproduction of Juanilama
(Lippia Alba) in the Caribbean Region of Costa Rica.” Adv.
Rafael Ocampo. Colorado. 1999. 20pp.
I conducted an integrated study of ethnofarmacology and biology
of a medicinal plant Lippia alba during the months of March
and April. The study was done in the Agro-ecological garden of Bougainvillea
S.A. and in its surrounding communities of Matina County, Limon
Province, Costa Rica. Juanilama is the common name of Lippia
alba in Costa Rica. My first goal was to determine the knowledge
of the communities about the Juanilama concentrating on its traditional
usage and if in fact it was used by interviewing elementary school
students and their parents. Juanilama continues to be one of the
well known plant, but the knowledge of its traditional uses has
decreased. Education and revaluation of this cultural aspect is
in need. Continuous extraction of Juanilama from the wild and inability
to adequately domesticate Juanilama puts it in danger of disappearance.
In result diminishes people using it and benefitting from its medicinal
properties. I have performed numerous propagation tests using asexual
means of reproduction by cuttings of branches of the plant. The
result of these tests showed the most economic and effective method
of reproduction of Juanilama in the Caribbean region of Costa Rica.
Strauss,
Ben, "Measuring the Effects of Syzygium jambos (Rosy
Apple, Myrtaceae) on Patterns of Understory Plant Diversity
in a Tropical Premontane Forest." Advisor: Gerardo Ávalos. Knox.
2005. 20pp.
The exotic tree species, Syzygium jambos (Rosy Apple, Myrtaceae),
was studied at the Municipal Forest of Atenas, Alajuela, Costa Rica,
from February 28 to April 22 of 2005. The objective of the study
was to determine what effect S. jambos is having on understory
other understory species is indicative of the invasiveness and competitiveness
of the exotic. No relationship was observed between the density
of the plant and canopy cover data or distance from the edge of
the forest, signifying a fairly homogenous forest that has already
been substantially colonized by the invader throughout. Data indicates
that S. jambos is having a negative effect on understory
diversity and so advice concerning the removal and control of the
species is given. Mechanical or biological controls are the best
bets for this forest fragment, given sufficient research, funds
and labor force to ensure a successful control program. Also discussed
is the forest's status as an area of protection for drinking water
of neighboring communities. Extirpation of exotics should be considered
as long as water protection is not affected.
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Wetzler,
Erica, "A Study of the Successional Process in Clear Cut Areas
of El Carmen de Parrita, Costa Rica." Adv. Alvaro Castaing. St.
Olaf College. 1995. 40pp.
The successional process was investigated at three sites of land
near El Carmen de Parrita, Costa Rica. Each site was divided into
quadrants and samples of each species within random quadrants were
collected. Height and leaf area measurements were also recorded
for each plant. Results indicated that while diversity, frequency,
and density of scientific families are directly related to the number
of years since an area had been cut, leaf area is more closely related
to changes in altitude. Temperature readings were recorded as well.
Results demonstrated that temperature was inversely related to altitude
and directly related to proximity to the native forest.
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